Ray Romano Manny (voice) John Leguizamo Sid (voice) Denis Leary Diego (voice) Seann William Scott Crash (voice) Josh Peck Eddie (voice) Queen Latifah Ellie (voice) Jennifer Lopez Shira (voice) Peter Dinklage Captain Gutt (voice) Wanda Sykes Granny (voice) Keke Palmer Peaches (voice) Josh Gad Louis (voice) Nick Frost Flynn (voice) Aziz Ansari Squint (voice) Alain Chabat Silas (voice) Kunal Nayyar Gupta (voice) Drake Ethan (voice) Nicki Minaj Steffie (voice) Rebel Wilson Raz (voice) Alan Tudyk Milton / Hunky Siren (voice) Ester Dean Female Siren (voice) Eddie 'Piolin' Sotelo Uncle Fungus (voice) Joy Behar Eunice (voice) Heather Morris Katie (voice) Chris Wedge Scrat (voice) Karen Disher Scratte (voice) Marieve Herington Additional Voices (voice) Ben Gleib Marshall (voice) Simon Pegg Buck (voice) Patrick Stewart Ariscratle (voice) Edita Brychta Additional Voices (voice) Dennis T. Carnegie Additional Voices (voice) Aimée Castle Additional Voices (voice) Jim Conroy Additional Voices (voice) Ariel Winter Additional Voices (voice) Fred Tatasciore Additional Voices (voice)